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Test Info:

<!doctype html>
<meta charset="utf-8">
<link rel="author" href="mailto:emilio@crisal.io" title="Emilio Cobos Álvarez">
<link rel="author" href="https://mozilla.org" title="Mozilla">
<link rel="match" href="part-after-combinator-invalidation-ref.html">
<style>
.inactive > ::part(content) {
color: red;
}
</style>
<div class="inactive">
<div id="host">
<template shadowrootmode="open">
<div part="content">What color am I?</div>
</template>
</div>
</div>
<script>
onload = () => {
host.getBoundingClientRect();
host.parentNode.className = "";
host.getBoundingClientRect();
};
</script>