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Test Info:

<!DOCTYPE html>
<script src="/resources/testharness.js"></script>
<script src="/resources/testharnessreport.js"></script>
<style>
@keyframes test {
to { --x: to; }
}
#target {
--x: underlying;
animation: test 1s;
}
</style>
<div id="target"></div>
<script>
CSS.registerProperty({
name: '--x',
syntax: '*',
inherits: false,
});
test(() => {
target.style.animationDelay = '-0.25s';
assert_equals(getComputedStyle(target).getPropertyValue('--x'), 'underlying', 'at 25%');
target.style.animationDelay = '-0.75s';
assert_equals(getComputedStyle(target).getPropertyValue('--x'), 'to', 'at 25%');
}, 'CSS Animations neutral keyframes on registered custom properties should produce the underlying value');
</script>